بررسی صفات افتراقی ساختار تشریحی برگ در تاکسونومی هفت گونه Alopecurus (Poaceae) در ایران
محورهای موضوعی : ژنتیکمحبوبه خطابخش 1 , مصطفی اسدی 2 , مجتبی خیام نکویی 3
1 - دانشگاه آزاد اسلامی واحد علوم و تحقیقات تهران
2 - عضو هیئت علمی موسسه تحقیقات جنگلها و مراتع تهران
3 - ریاست پژوهشکده بیوتکنولوژی کشاورزی اصفهان
کلید واژه: ساختار تشریحی برگ, Alopecurus,
چکیده مقاله :
این مطالعه امکان بررسی دسته بندیهای طبیعی گونه‌های Alopecurus را بر اساس ویژگیهای تشریحی برگ فراهم می‌کند. در ایران تاکسونومی این جنس بسیار اندک بوده و ساختار تشریحی برگ در تاکسونومی آن استفاده نشده است. نمونه های7 گونه شناسایی شده از لحاظ ریختی بررسی شدند و صفات ریختی افتراقی مشخص شدند. همچنین نمونه ها از لحاظ ساختار تشریحی برگ در برش عرضی و اپیدرمها مورد بررسی قرار گرفتند و صفات تشریحی برگ مشخص شدند. صفات تشریحی افتراقی متمایز شدند. 38 صفت ریختی و 18 صفت تشریحی به عنوان صفات افتراقی بین گونه‌ای از لحاظ تاکسونومی عددی بطور جداگانه و مجموعا بصورت 56 صفت بررسی شدند. نتایج این تحقیق دسته بندیهای طبیعی را تایید کردند.
This study provides the possibility of investigating the natural classifications of Alopecurus species based on leaf anatomical features. In Iran, the taxonomy of this genus is very small and the anatomical structure of the leaf has not been used in its taxonomy. The samples of 7 identified species were examined in terms of shape and differential shape characteristics were determined. Also, the samples were examined in terms of the anatomical structure of the leaf in transverse section and epidermis, and the anatomical characteristics of the leaf were determined. Differential anatomical features were distinguished. 38 morphological traits and 18 anatomical traits were investigated as differentiating traits between species in terms of numerical taxonomy separately and in total 56 traits. The results of this research confirmed the natural classifications.
Blinnikov, M., 2002. Phytolith Studies in Western North America. Botanical electronic News 282: 1188- 1193.
Bor, N.L., 1970. Gramineae, In: Rechinger, K. H., Flora Iranica, No.70. Akademische Druech-u, Verlagsanstalt, Graz, Austria, P: 275-285.
Bor, N.L., 1968. Graminae, In flora of Iraq, Vol. 9. Mministry of Agriculture of the Republic of Iraq, Baghdad, P: 282-295.
Clayton, W.D. and S.A. Renvoize, 1986. Genera Graminum Grasses of the Word. HER Majesty's Stationery Office, London.
Davila, P. and L.G. Clark, 1990. Scanning Electron Microscopy Survey of Leaf Epidermis of Sorghastrum (Poaceae). Amer. J. Bot. 77(4): 499-511.
Davis, P.H., R.R. Mill and K. Tan, 1985. Flora of Turkey and the East Aegean Islands, Vol.9. Edinburgh, P: 373-387.
Dogan, M., 1999. A Concise Taxonomic Revision of the Genus Alopecurus. Tr. J. Bot. 23: 245-262.
Hubbard, C.E., 1954. Grasses, A guide to their Structure, Identification, Uses, and Distribution in the British Isles. Penguin Books.
Metcalf, C.R., 1960. Anatomy of the Monocotyledons I. Gramineae. Oxford, P: 18-21.
Metcalf, C.R., 1963. Comparative Anatomy as a Modern Botanical Discipline with Special Reference to Recent Advances in the Systematic of Monocotyledons. Adv. Bot. Res. 1: 101-147.
Naik, V.N., 1984. Taxonomy of Angiosperms. TATA Mcgraw-Hill, Delhi, P: 128-156.
Parry, D. W. and F. Smithson, 1964. Type of Opalin Silica Depositions in the Leaves of British Grasses. Annales of Bot. N. S. 28(109): 169-185.
Rozhevits, R. Yu. and B. K. Shishkin, 1963. Flora of the U.S.S.R., Vol. II (Gramineae). The National Science Foundation, Washington, P: 111-129.
Stancik, D., 2004. F. dinirica and F. guaramacalana (Poaceae), Tow New Species from the Venezuelan Andes. Novon(14) 3: 341-344.
Tzvelev, N. N., 1983. Grasses of the Soviet Union, Part,I. Oxonian Press, PVT. LTD, P: 548-566.
Vieira, R.C., D.M.S. Gomes, L.S. Sarahyba and R.C.O. Arruda, 2002. Leaf Anatomy of three Herbaceous Bamboo Species. Braz. J. Biol. Vol.62 no 4b, 1519-6984.
Whang, S.S., K. Kim and W.M. Hess, 1998. Variation of Silica Bodies in Leaf Epidermal Long Cells Within and Among Seventeen Species of Oryza (Poaceae). Amer. J. Bot. 85(4): 461-466.
_||_Blinnikov, M., 2002. Phytolith Studies in Western North America. Botanical electronic News 282: 1188- 1193.
Bor, N.L., 1970. Gramineae, In: Rechinger, K. H., Flora Iranica, No.70. Akademische Druech-u, Verlagsanstalt, Graz, Austria, P: 275-285.
Bor, N.L., 1968. Graminae, In flora of Iraq, Vol. 9. Mministry of Agriculture of the Republic of Iraq, Baghdad, P: 282-295.
Clayton, W.D. and S.A. Renvoize, 1986. Genera Graminum Grasses of the Word. HER Majesty's Stationery Office, London.
Davila, P. and L.G. Clark, 1990. Scanning Electron Microscopy Survey of Leaf Epidermis of Sorghastrum (Poaceae). Amer. J. Bot. 77(4): 499-511.
Davis, P.H., R.R. Mill and K. Tan, 1985. Flora of Turkey and the East Aegean Islands, Vol.9. Edinburgh, P: 373-387.
Dogan, M., 1999. A Concise Taxonomic Revision of the Genus Alopecurus. Tr. J. Bot. 23: 245-262.
Hubbard, C.E., 1954. Grasses, A guide to their Structure, Identification, Uses, and Distribution in the British Isles. Penguin Books.
Metcalf, C.R., 1960. Anatomy of the Monocotyledons I. Gramineae. Oxford, P: 18-21.
Metcalf, C.R., 1963. Comparative Anatomy as a Modern Botanical Discipline with Special Reference to Recent Advances in the Systematic of Monocotyledons. Adv. Bot. Res. 1: 101-147.
Naik, V.N., 1984. Taxonomy of Angiosperms. TATA Mcgraw-Hill, Delhi, P: 128-156.
Parry, D. W. and F. Smithson, 1964. Type of Opalin Silica Depositions in the Leaves of British Grasses. Annales of Bot. N. S. 28(109): 169-185.
Rozhevits, R. Yu. and B. K. Shishkin, 1963. Flora of the U.S.S.R., Vol. II (Gramineae). The National Science Foundation, Washington, P: 111-129.
Stancik, D., 2004. F. dinirica and F. guaramacalana (Poaceae), Tow New Species from the Venezuelan Andes. Novon(14) 3: 341-344.
Tzvelev, N. N., 1983. Grasses of the Soviet Union, Part,I. Oxonian Press, PVT. LTD, P: 548-566.
Vieira, R.C., D.M.S. Gomes, L.S. Sarahyba and R.C.O. Arruda, 2002. Leaf Anatomy of three Herbaceous Bamboo Species. Braz. J. Biol. Vol.62 no 4b, 1519-6984.
Whang, S.S., K. Kim and W.M. Hess, 1998. Variation of Silica Bodies in Leaf Epidermal Long Cells Within and Among Seventeen Species of Oryza (Poaceae). Amer. J. Bot. 85(4): 461-466.