الگوی پراکنش مکانی و حجم خشکه دارها در راشستان های شمال ایران (مطالعه موردی: بخش گرازبن جنگل خیرود)
محورهای موضوعی : جنگلداریزهرا نوری 1 , جهانگیر فقهی 2 , محمدرضا مروی مهاجر 3
1 - دانش آموخته دکتری جنگلداری، دانشگاه تهران، دانشکده منابع طبیعی، گروه جنگلداری و اقتصاد جنگل، کرج، ایران
2 - دانشیار گروه جنگلداری و اقتصاد جنگل، دانشگاه تهران، دانشکده منابع طبیعی،کرج، ایران
3 - استاد گروه جنگلداری و اقتصاد جنگل، دانشگاه تهران، دانشکده منابع طبیعی،کرج، ایران
کلید واژه: خشکهدار افتاده, خشکهدار سرپا, تابع K-Ripley, درجات پوسیدگی, جنگل راش, جنگلهای خزری,
چکیده مقاله :
پژوهش حاضر به مطالعه و کمی نمودن برخی از مهمترین خصوصیات خشکه دارها از جمله الگوی پراکنش مکانی، حجم و مراحل پوسیدگی آنها در یک جنگل راش می پردازد. به این منظور پارسل 319 بخش گرازبن جنگل آموزشی و پژوهشی خیرود به مساحت 48 هکتار که تا زمان آماربرداری در آن هیچ گونه عملیات مدیریتی در قالب طرح های جنگلداری انجام نگرفته به عنوان منطقه مورد مطالعه انتخاب شد و گونه، قطر برابر سینه و ارتفاع خشکه دارهای سرپا مورد اندازه گیری قرار گرفت. در خشکه دارهای افتاده نیز گونه درختی، قطر میانی و طول آنها ثبت شد. همچنین به منظور مطالعه الگوی مکانی خشکه دارها، موقعیت هر یک از خشکه دارها با روش آزیموت- فاصله و با استفاده از دستگاه فاصله یاب لیزری بدست آمد. نتایج مطالعه حاضر نشان داد حجم در هکتار خشکه دارها در منطقه مورد مطالعه 93/34 متر مکعب در هکتار میباشد که حجم خشکه دارهای افتاده و سرپا به ترتیب 835/28 و 622/6 مترمکعب در هکتار است. بیشترین درصد خشکه دارها متعلق به درجه پوسیدگی نرم درجه یک میباشد که 44 درصد از کل حجم خشکه دارها را به خود اختصاص داده است. استفاده از تابع K-Ripley نشان داد الگوی پراکنش خشکه دارهای افتاده، سرپا و همچنین کل خشکه دارهای موجود در منطقه مورد مطالعه کپه ای می باشد.
The aim of this study is to analyze the spatial pattern, volume and decay stages of dead woods in beech forests, Northern Iran. The current research was carried out in compartment 319, Gorazbon District, Khayroud Forests dominated by Fagus orientalis in which no management practices have been done before this inventory. Dead trees characteristics including species, DBH and height of snags and mid-trunk diameter, length and decay grade of downed trees were recorded. The coordinates of all dead trees (snags and downed) were recorded using azimuth-distance method by laser device. The results showed that the volume of dead trees in study area is 34.93 m3/ha that the volume of snags and downed dead trees is 6.622 and 28.835 m3/ha respectively. Decay class I has accounted for most of the volume of dead trees (44%). The spatial pattern of dead trees was determined as clustered, using K-Ripley function.
References
1- Aakala, T., T. Kuuluvainen, S. Gauthier & L. Grandpré, 2008. Standing dead trees and their decay-class dynamics in the north eastern boreal old-growth forests of Quebec. Forest Ecology and Managment 255: 410–420.
2- Akhavan, R., Kh. Sagheb Talebi, M. Hassani, & P. Parhizkar, 2010. Spatial patterns in untouched beech (Fagus orientalis Lipsky) stand over forest development stages in Kelardasht region of Iran. Iranian Journal of Forest and Poplar Research 18(2): 322-336.
3- Albrecht, L., 1991. Die Bedeutungg des toten Holzes im Wald Forstw.cbl.110: 106-113.
4- Amanzadeh B., Kh. Sagheb-Talebi, B. Sotoudeh Foumani, F. Fadaie, J. Camarero & J. Linares, 2013. Spatial Distribution and Volume of Dead Wood in Unmanaged Caspian Beech (Fagus orientalis) Forests from Northern Iran. Forests 4: 751-765.
5-Arevalo, J.R. & J.M. Fernandez-Palacios, 2003. Spatial patterns of trees and juveniles in a laurel forest of Tenerife, Canary Islands. Plant Ecology 165: 1–10.
6– Baker, TR. & K.J. Chaw, 2011. Manual for coarse woody debris measurement in rainforest plots. Natural environment research council. 8 p.
7 - Besag, J. 1977. Contribution to the discussion of Dr. Ripley's paper. Journal of the Royal Statistical Society. 39(2): 193-195.
8- Bohl, J. & B. Brandli, 2007. Deadwood volume assessment in third Swiss National Forest Inventory: methods and first results. European Journal of Forest Research 126(3):449 – 457.
9- Brunet, J., O. Fritz, & G. Richnau, 2010. Biodiversity in European beech forests—A review with recommendations for sustainable forest management. Ecology Bulletine 53: 77–94.
10-Christensen, M. & J. Clausen, 2005. Managing dead wood in European beech forests, nat_man (nature –based-management of beech in Europe) project.
11- Christensen, M., K.P. Hahn, E. Mountford & M. Odor, 2005. Dead wood in European beech (Fagus sylvatica) forest reserves. Forest Ecology and management 210: 267–282.
12- Cressie, N.A.C., 1993. Statistics for spatial data.Wiley, NewYork, 900 pp.
13- Diggle, P.J., 2003. Statistical Analysis of Point Patterns. Arnold, London, 455 pp.
14- Duncan, R.P., 1991. Competition and the coexistence of species in a mixed podocarp stand. Journal of Ecology 79: 1073–1084.
15- Etemad, V., 2009. Optimal management of kheirud forest: Forest typology map. Iran national science foundation, 120 pp.
16- Gossner, M.M., T. Lachat, J. Brunet, G. Isacsson, C. Bouget, H. Brustel, R. Brandl, W.W. Weisser & J. Müller, 2013. Current near-to-nature forest management effects on functional trait composition of saproxylic beetles in Beech Forests. Conservation Biology 27: 605–614.
17_ Habashi, H., 1997. Syudying the importance of dead trees in Vaz forest of Mazandaran, Msc Thesis, Forestry, natural resources faculty, Tarbiat modarres University.
18- Harmon, M.E., J.F. Franklin, F.J. Swanson, P. Sollins, S.V. Gregory, J.D. Lattin, N.H. Anderson, S.P. Cline, N.G. Aumen, & J.R. Sedell, 1986. Ecology of coarse woody debris in temperate ecosystems. Advances in Ecological Researc 15: 133–276.
19- Heilmann-Clausen, J., & M. Christensen, 2004. Does size matter? On the importance of various dead wood fractions for fungal diversity in Danish beech forests. Forest Ecology and management 201: 105–117.
20-Kirby, K.J., C.M. Reid, R.C. Thomas, & F.B. Goldsmith, 2007. Preliminary estimates of fallen dead wood and standing dead trees in managed and unmanaged forests in Britain. Journal of Applied Ecology 35: 148–155.
21- Kooch, Y., S. M. Hosseini, M. Akbarinia, M. Tabari & S. Gh. Jalali, 2010. The role of dead tree in regeneration density of mixed beech stand (case study: Sardabrood forests, Chalous, Mazindaran), Iranian Journal of Forest 2(2): 93-103.
22- Lanna, J. & P. Laroque, 2007. Decay progression and classification in two old-growth forests in Atlantic Canada, Forest Ecology and Management 238: 293 - 301.
23- Lassauce, A., Y. Paillet, H. Jactel, & C. Bouget, 2011. Deadwood as a surrogate for forest biodiversity: Meta-Analysis of correlations between deadwood volume and species richness of saproxylic organisms. Ecological Indicator 11: 1027–1039.
24- Lorimer, C.G., & A.S. White, 2003. Scale and frequency of natural disturbances in the Northeastern United States: Implications for early successional forest habitats and regional age distributions. Forest Ecology and Managment 185: 41–64.
25- Marvie Mohadjer, M.R. 2005. Silviculture. University of Tehran Press, Tehran, 387 pp.
26- Marvie Mohadjer, M. R., M. Zobeiri, V. Etemad & M. Jourgholami, 2008. Performing of the single selection method at compartment level and necessity for full inventory of tree species (Case study: Gorazbon District in Kheyrud Forest, North of Iran). Journal of natural resources of Iran 61(4): 889-908.
27- Mohammad Nejad kiasari, Sh. & R. Rahmani, 2001. Effect of dead trees on the frequency of natural regeneration in a beech-hornbeam forest (Jamaleddinkola, Mazandaran). Journal of the Iranian natural resources 54(2):143-151.
28-Nouri, Z., 2012. Multi-purpose Inventory Method Based on Ecological Approach for Beech Forests of Iran (Case study: Gorazbon district, Kheyrud forest). PhD thesis, natural resources faculty, Tehran University.
29- Passovoy, M., & Z. Fule, 2006. Snag and woody debris dynamics following severe wildfires in northern Arizona ponderosa pine forests, Forest Ecology and Management 223: 237-246.
30- Russell, M.B., L.S. Kenefic, A.R. Weiskittel, J.J. Puhlick, & J.C. Brissette, 2012. Assessing and modeling standing deadwood attributes under alternative silvicultural regimes in the Acadian Forest region of Maine, USA. Canadian Journal of Forest Research. 42: 1873–1883.
31– Santiago, J.M., & D.R. Amanda, 2005. Dead trees resources for forest wildlife, Extension fact sheet, Ohio University Express, Ohio.
32- Sefidi, K., 2007. Quantitative and qualitative studying of dead trees in a managed forest (case study: Patom and Namkhaneh District of Kheirud forest), Msc thesis, natural resources faculty, Tehran University.
33- Sefidi, K. & M. R. Marvie Mohadjer, 2009. Amount and quality of dead trees (snag and logs) in a mixed beech forest with different management histories, Journal of Forest and Wood Products 62(2): 191-202.
34- Siitonen, J., P. Martikainen, P. Punttila & J. Rauh, 2000. Coarse woody debris and stand characteristics in mature managed and old – growth boreal mesic forests in southern Finland, Forest Ecology and Management 128: 211-225.
35- Wisdom, M.J. & L.J. Bate, 2008. Snag density varies with intensity of timber harvest and human access. Forest Ecology and Management 255: 2085–2093.
36- Zolfeghari, E., 2004. Ecological and Silvicultural investigation of dead trees in beech forests of Iran, Msc Thesis, natural resources faculty, Tehran University.
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