تعیین مناسبترین سطح قطعه نمونه به منظور بررسی الگوی مکانی درختان ارس (Juniperus excelsa) (مطالعه موردی: منطقه آتشگاه کرج)
محورهای موضوعی : منابع طبیعیوحید علی جانی 1 , سید محمد معین صادقی 2 , منوچهر نمیرانیان 3 , رضا اخوان 4
1 - دکتری جنگلداری و اقتصاد جنگل، دانشکده منابع طبیعی، دانشگاه تهران، تهران، ایران.* (مسوول مکاتبات)
2 - دکتری جنگلشناسی و اکولوژی جنگل، دانشکده منابع طبیعی، دانشگاه تهران، تهران، ایران.
3 - استاد گروه جنگلداری و اقتصاد جنگل، دانشکده منابع طبیعی، دانشگاه تهران، تهران، ایران.
4 - دانشیار پژوهش، مؤسسه تحقیقات جنگلها و مراتع کشور، سازمان تحقیقات، آموزش و ترویج کشاورزی، تهران، ایران.
کلید واژه: رقابت, تابع g(r), روابط بین گونهای, ایران-تورانی,
چکیده مقاله :
زمینه و هدف: اخیراً در تحلیل های مرتبط با الگوی مکانی درختان، روش های نوینی توسعه یافته اند که به منظور بررسی، به یک نقشه کامل از موقعیت مکانی درختان نیاز دارند. به کارگیری این روش ها به مقیاس مطالعه بستگی دارد به نحوی که تغییر مقیاس مطالعه می تواند سبب تغییراتی در الگوی مکانی درختان شود. پژوهش حاضر با هدف تعیین مناسب ترین سطح قطعهنمونه جهت کمیسازی الگوی مکانی درختان ارس دامنه جنوبی البرز صورت پذیرفت. روش بررسی: بدین منظور قطعهنمونه ای به مساحت شش هکتار انتخاب و الگوی مکانی درختان با استفاده از تابع g(r) مورد بررسی قرار گرفت. سپس سطح این قطعهنمونه به پنج، چهار، سه، دو و یک هکتار کاهش و الگوی مکانی درختان با استفاده از تابع g(r) بررسی و نسبت به سطح مبنای شش هکتاری مقایسه شد. یافته ها: نتایج حاصل از این تابع نشاندهنده ی الگوی مکانی یکنواخت در فواصل بین درختی تا چهار متر، الگوی کپه ای در فواصل شش تا ده متر و الگوی تصادفی در سایر فواصل می باشد. نتایج حاصل از مقایسه نمودارهای تابع g(r) در مساحتهای مختلف نشان داد که روند کلی تابع g(r) تا سطح سه هکتار تقریباً ثابت و در سطوح کوچک تر، تغییر می کند، بنابراین حداقل سطح مناسب در این بررسی، سه هکتار به دست آمد. بحث و نتیجه گیری: تعیین مناسبترین سطح قطعه نمونه در مطالعات ساختاری ضمن افزایش دقت مطالعات می تواند سبب کاهش زمان و هزینه مطالعه گردد.
Background and Objective: In recent years, new methods have been developed for analyzing spatial pattern of trees, and for verifying the spatial pattern, they need to complete map of spatial location of these trees. The use of these methods depends on study scale in a way that changes in study scale cause changes in spatial pattern. In this regard, present study aims to determine the optimum area of sample plots to quantify the spatial pattern of Juniperus Excelsa M. Beib. In southern facing slopes of Alborz mountains. Method: A plot with area of six hectares was selected and spatial pattern of trees were evaluated by using g(r) function. In next step, area of plot decreased to five, four, three, two, and one hectare and spatial pattern of trees were determined by using g(r) function and were compared with six hectare plot. Findings: The results of this function indicates regular spatial pattern in trees distances to four meter, cluster pattern at distances of six to ten meter, and random pattern shows in other distances. The results of comparing figures of g(r) functions in different area showed that the overall trend of this functions almost constant up to three hectares. Hence, in this study the appropriate area is achieved three hectares. Discussion and Conclusion: It should be noted that in structure studies to determine the best area of plot could reduce time and costs while increasing the precision of the study.
- Ali Ahmad Korori , S., Khoshnevis, M., Matinizadeh, M., 2010. Comprehensive studies of Juniperus species in Iran, Pooneh publication. (In Persian)
- Ahmed, M., Husain, T., Sheikh, A.H., Sadruddin, S., Siddiqui, H.M.F., 2006. Phytosociology and structure of Himalayan forests from different climatic zones of Pakistan, Pakistan Journal of Botany, Vol. 38(2), pp. 361-383.
- Mohamadnezhad Kiasari, SH., Safaei, M., Nourouzi, Sh. A., Ahmadian, S. H., Mataji, A. 2009. The evaluation of protection along with water spreading operations on the quantitative improvement of Greek Juniper seedlings (Juniperis excelsa Beib.) (Case study: Mazandaran-Poshtkoh watershed), Water and soil science (Journal of science and technology of agriculture and natural), Vol. 13(48), pp. 415-425. (In Persian)
- Livingston, R.B., 1972. Influence of birds, stone and soil on the establishment of pasture juniper, Juniper communis and Red cedar and J. virginiana in New England pastures, Ecology, Vol. 53(6), pp. 1141–1147.
- Krebs, C. J., 1999. Ecological Methodology, 2nd ed., Addison Welsey Educational Publication. Inc., USA.
- Marie-Agnes, M., Audrey, R., 2003. A model to assess relationships between forest dynamics and spatial structure. Journal of Vegetation Science, Vol. 14(6), pp. 823-834.
- Pommerening, A., Stoyan, D., 2008. Reconstructing spatial tree point patterns from nearest neighbour summary statistics measured in small subwindows. Canadian Journal of Forest Research, Vol. 38, pp. 1110–1122.
- Sohrabi, H., 2014. Spatial pattern of woody species in Chartagh forest reserve, Ardal. Iranian Journal of Forest and Poplar Research Vol. 22(1), pp.27-38. (In Persian)
- Akhavan, R., Sagheb-Talebi, Kh., Hassani, M., Parhizkar, P., 2010. Spatial patterns in untouched beech (Fagus orientalis Lipsky) stands over forest development stages in Kelardasht region of Iran. Iranian Journal of Forest and Poplar Research, Vol. 18(2), pp. 322-336. (In Persian)
- Li, F., Zhang, L., 2007. Comparisons of point pattern analysis methods for classifying the spatial distributions of spruce-fir stands in the north-east USA. Journal of Forestry, Vol 3, pp. 337-349.
- Batoubeh, P., Akhavan, R., Pourhashemi, M., Kia-Daliri, H., 2013. Determining the minimum plot size to study the spatial patterns of manna oak trees (Quercus brantii Lindl.) using Ripley,s K-function at less-disturbed stands in Marivan forests. Journal of Forest and Wood Product (Iranian Journal of Natural Resources), Vol. 66(1), pp. 27-38. (In Persian)
- Erfanifard, Y., Rezayan, F., 2014. Suitable Methods in Spatial Pattern Analysis of Heterogeneous Wild Pistachio (Pistacia atlantica Desf.) Woodlands in Zagros, Iran. Iranian Journal of Applied Ecology, Vol. 3(9), pp. 81-91. (In Persian)
- Illian, J., Penttinen, A., Stoyan, H., Stoyan, D., 2008. Statistical Analysis and Modelling of Spatial Point Patterns. John Wiley & Sons Inc, West Sussex.
- Getzin, S., Dean, C., He, F., Trofymow, J., Wiegand, K., Wiegend, T., 2006. Spatial patterns and competition of tree species in a Douglas-fir chronosequence on Vancouver Island, Ecography, Vol. 29, pp. 671–682.
- Hosseinzadeh, J., Namiranian, M., Marvi Mohajer, M., Zahedi Amiri, Gh., 2004. Structure of less degraded oak forests in Ilam province. Iranian Journal of Natural Resources. Vol. 57(1), pp. 75-90. (In Persian)
- Haidari, M., Namiranian, M., Zobiri, M., Gahramany, L., 2014. Investigation on appropriate inventory method for determining structure of Northern Zagros Forests (Case study: Blake Forests, Baneh). Iranian Journal of Forest and Poplar Research, Vol. 21(3), pp. 467-480. (In Persian)
- Fallah, A., 2000. Investigation of the natural beech (Fagus orientalis) stands structure in Mazandaran and Golestan province. Unpublished Ph.D thesis, University of Tehran, Faculty of Natural Resources. 202 p. (In Persian)
- Sagheb Talebi, Kh., Ghorchi Beigi, K., Eslami, A., Shahnavazi, H., Mousavi Mir Kelayee, S., 2001. Structure of Caspian beech forests and the potential of applying selection system in them, Proceeding Second International Conference of Forest and Industrial, Vol. 1, pp. 107-138. (In Persian)
- Sagheb Talebi, Kh., Schütz, J.Ph., 2002. The structure of natural oriental beech (Fagusorientalis) in the Caspian region of Iran and potential for the application of the group selection system. Forestry, Vol. 75(4), pp. 465-472.
- Askari, Y., Soltani, A., Sohrabi, H., 2014. Evaluation of Spatial distribution pattern of tree and shrub species in a central Zagros (Case study: Chahartagh forest reserve). Iranian Journal of Forest and Poplar Research Vol. 22(2), pp. 175-187. (In Persian)
- Unknown., 2015. Studies of the southern slope of Alborz (Karaj and Savojbolagh province). Natural resource and watersher management office, Alborz. (In Persian)
- Erfanifard, Y., Mahdian, F., 2012. Comparative investigation on the methods of true spatial pattern analysis of trees in forests (Case study: Wild pistachio research forest, Fars province, Iran). Iranian Journal of Forest and Poplar Research, Vol. 20(1), pp. 62-73. (In Persian)
- Stoyan, D., Stoyan, H., 1994. Fractals, Random Shapes and Point Fields: Methods of Geometrical Statistics. John Wiley & Sons Inc, West Sussex.
- Fortin, M.J., Dale, M.R.T., 2005. Spatial Analysis. Cambridge University Press, Cambridge.
- Guo, Y., Lu, J., Franklin, S.B., Wang, Q., Zhang, Y., Xu, K., Bao, D., Qiao, X., Huang, H., Lu, Z., Jiang, M., 2013. Spatial distribution of tree species in a species-rich subtropical mountain forest in central China. Canadian Journal of Forest Research, Vol (43), pp. 826-835.
- Szmyt, J., 2014. Spatial statistics in ecological analysis: from indices to functions. Silva Fennica,Vol. 48(1), pp. 1-31.
- Mason, W. L., Connolly, T., Pommerening, A., Edwards, C., 2007. Spatial structure of semi-natural and plantation stands of Scots pine (Pinus sylvestris L.) in northern Scotland. Forestry. Vol. 80. Pp. 567-586.
- Getzin, S., Worbes, M., Wiegand, T., Wiegand, K., 2011. Size dominance regulates tree spacing more than competition within height classes in tropical Cameroon. Journal of Tropical Ecology, Vol. 27, pp. 93-102.
- Zhu, Y., Xiangcheng, X., Ren, H., Ma, K., 2010. Density dependence is prevalent in a heterogeneous subtropical forest. Oikos, Vol. (119), pp. 109-119.
- Strand, E.K., Robinson, A.P., Bunting, S.C., 2007. Spatial patterns on the sagebrush steppe/Western juniper ecotone. Plant Ecology, Vol. 190, pp.159-173.
- Bailey, T.C., Gatrell, A.C., 1998. Interactive spatial data analysis. Longman publication.
- Haase, P., 1995. Spatial pattern analysis in ecology based on Ripley’s K-function: introduction and methods of edge correction. Journal of Vegetation Science, Vol. 6, pp. 575-582.
- Martens, S.N., Breshears, D.D., Meyer, C.W., 1997. Scales of above-ground and below-ground competition in a semiarid woodland detected from spatial pattern. Journal of Vegetation Science, Vol. 8, pp. 655-664.
- Perry, G.L.W., 2004. SpPack: spatial point pattern analysis in Excel using visual basic for applications (VBA). Environmental Modelling and Software, Vol. 19, pp. 559-569.
- Schiffers, K., Schurr, F.M., Tielborger, K., Urbach, C., Moloney, K., Jeltsch, F., 2008. Dealing with virtual aggregation- a new index for analyzing heterogeneous point patterns. Journal of Ecography, Vol. 31, pp. 545-555.
- Beitollah, A., Pourmajidian, M.R., Sagheb Talebi, Kh., Hojjati, S.M., 2015. Spatial Pattern, competition and spatial association of trees in Mixed Hornbeam Stands using univariate and bivariate Ripley’s K-function (case study: Reserve area, district 3 of Asalem forests). Iranian Journal of Forest and Poplar Research, Vol. 23(1), pp. 37-52. (In Persian)
- Sadeghi1, S.M.M., Alijani, V., Namiranian, M., Mohamadizadeh, M. 2016. Structural characteristics of Juniperus excelsa in the mountainous forests of Alborz south facing slope (Case study: Atashgah, Karaj). Iranian Journal of Forest, Vol. 8(1), pp. 35-49. (In Persian)
- Holthuijzen, A.M.A., Sharik, T.L., 1985. Colonization of abandoned pastures by eastern red cedar (Juniperus virginiana L.), Canadian Journal of Forest Research, Vol. 15(6), pp. 1065-1068.
- Houle, G., Duchesne, M., 1999. The spatial pattern of a Juniperus communis var. depressa population on a continental dune in subarctic Québec, Canada. Canadian Journal of Forest Research, Vol. 29(4), pp. 446-450.
- Pielou, E.C., 1962. The use of plant-to-neighbour distances for the detection of competition, Journal of Ecology, Vol. 50, pp. 357-367.
- Antonovics, J., Levin, D.A., 1980. The ecological and genetic consequences of density-dependent regulation in plants, Annual Review of Ecology and Systematics, Vol. 11, pp. 411-452.
- Stoll, P., Bergius, E., 2005. Pattern and process: competition causes regular spacing of individuals within plant populations. Journal of Ecology, Vol. 93, pp. 395-403.
- Gray, L., He, F., 2009. Spatial point-pattern analysis for detecting density-dependent competition in a boreal chronosequence of Alberta. Forest Ecology and Management, Vol. 259, pp. 98-106.
- Trifković, S., 2013. Spatial distribution and composition of trees in an aging coppice forest in Japan. Formath, Vol. 1, pp. 35-53.
- Wiegand, T., Moloney, K.A., 2004. Rings, circles, and null-models for point pattern analysis in ecology. OIKOS, Vol. 104, pp. 209-229.
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- Ali Ahmad Korori , S., Khoshnevis, M., Matinizadeh, M., 2010. Comprehensive studies of Juniperus species in Iran, Pooneh publication. (In Persian)
- Ahmed, M., Husain, T., Sheikh, A.H., Sadruddin, S., Siddiqui, H.M.F., 2006. Phytosociology and structure of Himalayan forests from different climatic zones of Pakistan, Pakistan Journal of Botany, Vol. 38(2), pp. 361-383.
- Mohamadnezhad Kiasari, SH., Safaei, M., Nourouzi, Sh. A., Ahmadian, S. H., Mataji, A. 2009. The evaluation of protection along with water spreading operations on the quantitative improvement of Greek Juniper seedlings (Juniperis excelsa Beib.) (Case study: Mazandaran-Poshtkoh watershed), Water and soil science (Journal of science and technology of agriculture and natural), Vol. 13(48), pp. 415-425. (In Persian)
- Livingston, R.B., 1972. Influence of birds, stone and soil on the establishment of pasture juniper, Juniper communis and Red cedar and J. virginiana in New England pastures, Ecology, Vol. 53(6), pp. 1141–1147.
- Krebs, C. J., 1999. Ecological Methodology, 2nd ed., Addison Welsey Educational Publication. Inc., USA.
- Marie-Agnes, M., Audrey, R., 2003. A model to assess relationships between forest dynamics and spatial structure. Journal of Vegetation Science, Vol. 14(6), pp. 823-834.
- Pommerening, A., Stoyan, D., 2008. Reconstructing spatial tree point patterns from nearest neighbour summary statistics measured in small subwindows. Canadian Journal of Forest Research, Vol. 38, pp. 1110–1122.
- Sohrabi, H., 2014. Spatial pattern of woody species in Chartagh forest reserve, Ardal. Iranian Journal of Forest and Poplar Research Vol. 22(1), pp.27-38. (In Persian)
- Akhavan, R., Sagheb-Talebi, Kh., Hassani, M., Parhizkar, P., 2010. Spatial patterns in untouched beech (Fagus orientalis Lipsky) stands over forest development stages in Kelardasht region of Iran. Iranian Journal of Forest and Poplar Research, Vol. 18(2), pp. 322-336. (In Persian)
- Li, F., Zhang, L., 2007. Comparisons of point pattern analysis methods for classifying the spatial distributions of spruce-fir stands in the north-east USA. Journal of Forestry, Vol 3, pp. 337-349.
- Batoubeh, P., Akhavan, R., Pourhashemi, M., Kia-Daliri, H., 2013. Determining the minimum plot size to study the spatial patterns of manna oak trees (Quercus brantii Lindl.) using Ripley,s K-function at less-disturbed stands in Marivan forests. Journal of Forest and Wood Product (Iranian Journal of Natural Resources), Vol. 66(1), pp. 27-38. (In Persian)
- Erfanifard, Y., Rezayan, F., 2014. Suitable Methods in Spatial Pattern Analysis of Heterogeneous Wild Pistachio (Pistacia atlantica Desf.) Woodlands in Zagros, Iran. Iranian Journal of Applied Ecology, Vol. 3(9), pp. 81-91. (In Persian)
- Illian, J., Penttinen, A., Stoyan, H., Stoyan, D., 2008. Statistical Analysis and Modelling of Spatial Point Patterns. John Wiley & Sons Inc, West Sussex.
- Getzin, S., Dean, C., He, F., Trofymow, J., Wiegand, K., Wiegend, T., 2006. Spatial patterns and competition of tree species in a Douglas-fir chronosequence on Vancouver Island, Ecography, Vol. 29, pp. 671–682.
- Hosseinzadeh, J., Namiranian, M., Marvi Mohajer, M., Zahedi Amiri, Gh., 2004. Structure of less degraded oak forests in Ilam province. Iranian Journal of Natural Resources. Vol. 57(1), pp. 75-90. (In Persian)
- Haidari, M., Namiranian, M., Zobiri, M., Gahramany, L., 2014. Investigation on appropriate inventory method for determining structure of Northern Zagros Forests (Case study: Blake Forests, Baneh). Iranian Journal of Forest and Poplar Research, Vol. 21(3), pp. 467-480. (In Persian)
- Fallah, A., 2000. Investigation of the natural beech (Fagus orientalis) stands structure in Mazandaran and Golestan province. Unpublished Ph.D thesis, University of Tehran, Faculty of Natural Resources. 202 p. (In Persian)
- Sagheb Talebi, Kh., Ghorchi Beigi, K., Eslami, A., Shahnavazi, H., Mousavi Mir Kelayee, S., 2001. Structure of Caspian beech forests and the potential of applying selection system in them, Proceeding Second International Conference of Forest and Industrial, Vol. 1, pp. 107-138. (In Persian)
- Sagheb Talebi, Kh., Schütz, J.Ph., 2002. The structure of natural oriental beech (Fagusorientalis) in the Caspian region of Iran and potential for the application of the group selection system. Forestry, Vol. 75(4), pp. 465-472.
- Askari, Y., Soltani, A., Sohrabi, H., 2014. Evaluation of Spatial distribution pattern of tree and shrub species in a central Zagros (Case study: Chahartagh forest reserve). Iranian Journal of Forest and Poplar Research Vol. 22(2), pp. 175-187. (In Persian)
- Unknown., 2015. Studies of the southern slope of Alborz (Karaj and Savojbolagh province). Natural resource and watersher management office, Alborz. (In Persian)
- Erfanifard, Y., Mahdian, F., 2012. Comparative investigation on the methods of true spatial pattern analysis of trees in forests (Case study: Wild pistachio research forest, Fars province, Iran). Iranian Journal of Forest and Poplar Research, Vol. 20(1), pp. 62-73. (In Persian)
- Stoyan, D., Stoyan, H., 1994. Fractals, Random Shapes and Point Fields: Methods of Geometrical Statistics. John Wiley & Sons Inc, West Sussex.
- Fortin, M.J., Dale, M.R.T., 2005. Spatial Analysis. Cambridge University Press, Cambridge.
- Guo, Y., Lu, J., Franklin, S.B., Wang, Q., Zhang, Y., Xu, K., Bao, D., Qiao, X., Huang, H., Lu, Z., Jiang, M., 2013. Spatial distribution of tree species in a species-rich subtropical mountain forest in central China. Canadian Journal of Forest Research, Vol (43), pp. 826-835.
- Szmyt, J., 2014. Spatial statistics in ecological analysis: from indices to functions. Silva Fennica,Vol. 48(1), pp. 1-31.
- Mason, W. L., Connolly, T., Pommerening, A., Edwards, C., 2007. Spatial structure of semi-natural and plantation stands of Scots pine (Pinus sylvestris L.) in northern Scotland. Forestry. Vol. 80. Pp. 567-586.
- Getzin, S., Worbes, M., Wiegand, T., Wiegand, K., 2011. Size dominance regulates tree spacing more than competition within height classes in tropical Cameroon. Journal of Tropical Ecology, Vol. 27, pp. 93-102.
- Zhu, Y., Xiangcheng, X., Ren, H., Ma, K., 2010. Density dependence is prevalent in a heterogeneous subtropical forest. Oikos, Vol. (119), pp. 109-119.
- Strand, E.K., Robinson, A.P., Bunting, S.C., 2007. Spatial patterns on the sagebrush steppe/Western juniper ecotone. Plant Ecology, Vol. 190, pp.159-173.
- Bailey, T.C., Gatrell, A.C., 1998. Interactive spatial data analysis. Longman publication.
- Haase, P., 1995. Spatial pattern analysis in ecology based on Ripley’s K-function: introduction and methods of edge correction. Journal of Vegetation Science, Vol. 6, pp. 575-582.
- Martens, S.N., Breshears, D.D., Meyer, C.W., 1997. Scales of above-ground and below-ground competition in a semiarid woodland detected from spatial pattern. Journal of Vegetation Science, Vol. 8, pp. 655-664.
- Perry, G.L.W., 2004. SpPack: spatial point pattern analysis in Excel using visual basic for applications (VBA). Environmental Modelling and Software, Vol. 19, pp. 559-569.
- Schiffers, K., Schurr, F.M., Tielborger, K., Urbach, C., Moloney, K., Jeltsch, F., 2008. Dealing with virtual aggregation- a new index for analyzing heterogeneous point patterns. Journal of Ecography, Vol. 31, pp. 545-555.
- Beitollah, A., Pourmajidian, M.R., Sagheb Talebi, Kh., Hojjati, S.M., 2015. Spatial Pattern, competition and spatial association of trees in Mixed Hornbeam Stands using univariate and bivariate Ripley’s K-function (case study: Reserve area, district 3 of Asalem forests). Iranian Journal of Forest and Poplar Research, Vol. 23(1), pp. 37-52. (In Persian)
- Sadeghi1, S.M.M., Alijani, V., Namiranian, M., Mohamadizadeh, M. 2016. Structural characteristics of Juniperus excelsa in the mountainous forests of Alborz south facing slope (Case study: Atashgah, Karaj). Iranian Journal of Forest, Vol. 8(1), pp. 35-49. (In Persian)
- Holthuijzen, A.M.A., Sharik, T.L., 1985. Colonization of abandoned pastures by eastern red cedar (Juniperus virginiana L.), Canadian Journal of Forest Research, Vol. 15(6), pp. 1065-1068.
- Houle, G., Duchesne, M., 1999. The spatial pattern of a Juniperus communis var. depressa population on a continental dune in subarctic Québec, Canada. Canadian Journal of Forest Research, Vol. 29(4), pp. 446-450.
- Pielou, E.C., 1962. The use of plant-to-neighbour distances for the detection of competition, Journal of Ecology, Vol. 50, pp. 357-367.
- Antonovics, J., Levin, D.A., 1980. The ecological and genetic consequences of density-dependent regulation in plants, Annual Review of Ecology and Systematics, Vol. 11, pp. 411-452.
- Stoll, P., Bergius, E., 2005. Pattern and process: competition causes regular spacing of individuals within plant populations. Journal of Ecology, Vol. 93, pp. 395-403.
- Gray, L., He, F., 2009. Spatial point-pattern analysis for detecting density-dependent competition in a boreal chronosequence of Alberta. Forest Ecology and Management, Vol. 259, pp. 98-106.
- Trifković, S., 2013. Spatial distribution and composition of trees in an aging coppice forest in Japan. Formath, Vol. 1, pp. 35-53.
- Wiegand, T., Moloney, K.A., 2004. Rings, circles, and null-models for point pattern analysis in ecology. OIKOS, Vol. 104, pp. 209-229.